Unpaired fins in fish. Organs of movement - fins

; their organs that regulate movement and position in water, and in some ( flying fish) - also planning in the air.

The fins are cartilaginous or bony rays (radials) with skin-epidermal coverings on top.

The main types of fish fins are dorsal, anal, caudal, pair of abdominal and pair of pectoral.
Some fish also have adipose fins(they lack fin rays), located between the dorsal and caudal fins.
The fins are driven by muscles.

Often different types fish fins are modified for example, males viviparous fish use the anal fin as an organ for mating (the main function of the anal fin is similar to the function of the dorsal fin - it is a keel when the fish moves); at gourami modified filamentous pelvic fins are special tentacles; highly developed pectoral fins allow some fish to jump out of the water.

The fins of fish actively participate in movement, balancing the body of the fish in the water. In this case, the motor moment begins from the caudal fin, which pushes forward with a sharp movement. The tail fin is a kind of propulsion device for the fish. The dorsal and anal fins balance the fish's body in the water.

Different species of fish have different numbers of dorsal fins.
Herring and carp-like have one dorsal fin mullet-like and perch-like- two, y codlike- three.
They can also be located differently: pike- displaced far back, at herring-like, carp-like- in the middle of the ridge, at perch and cod- closer to the head. U mackerel, tuna and saury there are small additional fins behind the dorsal and anal fins.

The pectoral fins are used by the fish when swimming slowly, and together with the pelvic and caudal fins they maintain the balance of the fish’s body in the water. Many bottom-dwelling fish move along the ground using pectoral fins.
However, in some fish ( moray eels, for example) pectoral and ventral fins are absent. Some species also lack a tail: gymnots, ramfichtids, seahorses, stingrays, sunfish and other species.

Three-spined stickleback

In general, the more developed a fish's fins, the more suited it is to swimming in calm water.

In addition to movement in water, air, on the ground; jumping, jumping, fins help different types of fish attach to the substrate (sucker fins in bulls), look for food ( triggles), have protective functions ( sticklebacks).
Some types of fish ( scorpionfish) have poisonous glands at the base of the spines of the dorsal fin. There are also fish without fins at all: cyclostomes.

Fins

organs of movement of aquatic animals. Among invertebrates, P. have pelagic forms of gastropods and cephalopods and setaceous-maxillary. U gastropods P. are a modified leg; in cephalopods, they are lateral folds of skin. The chaetomagnaths are characterized by lateral and caudal wings formed by folds of skin. Among modern vertebrates, cyclostomes, fish, some amphibians, and mammals have P. In cyclostomes there are only unpaired P.: anterior and posterior dorsal (in lampreys) and caudal.

In fish, there are paired and unpaired P. Paired ones are represented by anterior (thoracic) and posterior (abdominal) ones. In some fish, such as cod and blenny, the abdominal pectorals are sometimes located in front of the pectoral ones. The skeleton of paired limbs consists of cartilaginous or bone rays, which are attached to the skeleton of the limb girdles (See Limb girdles) ( rice. 1 ). The main function of paired propellers is the direction of fish movement in the vertical plane (depth rudders). In a number of fish, paired parasites perform the functions of active swimming organs (See Swimming) or are used for gliding in the air (in flying fish), crawling along the bottom, or moving on land (in fish that periodically leave the water, for example, in representatives of the tropical genus Periophthalmus , which, with the help of chest pectorals, can even climb trees). The skeleton of unpaired P. - dorsal (often divided into 2 and sometimes into 3 parts), anus (sometimes divided into 2 parts) and caudal - consists of cartilaginous or bone rays lying between the lateral muscles of the body ( rice. 2 ). The skeletal rays of the caudal vertebrae are connected to the posterior end of the spine (in some fish they are replaced by the spinous processes of the vertebrae).

The peripheral parts of the P. are supported by thin rays of horn-like or bone tissue. In spiny-finned fish, the anterior of these rays thicken and form hard spines, sometimes associated with poisonous glands. Muscles that stretch the lobe of the pancreas are attached to the base of these rays. The dorsal and anal parasites serve to regulate the direction of movement of the fish, but sometimes they can also be organs of forward movement or perform additional functions (for example, attracting prey). The caudal part, which varies greatly in shape in different fish, is the main organ of movement.

In the process of the evolution of vertebrates, the P. of fish probably arose from a continuous fold of skin that ran along the back of the animal, went around the rear end of its body and continued on the ventral side to the anus, then divided into two lateral folds that continued to the gill slits; This is the position of the fin folds in the modern primitive chordate - Lancelet a. It can be assumed that during the evolution of animals, skeletal elements formed in some places of such folds and in the intervals the folds disappeared, which led to the emergence of unpaired folds in cyclostomes and fish, and paired ones in fish. This is supported by the presence of lateral folds or venom of spines in the most ancient vertebrates (some jawless, acanthodia) and the fact that in modern fish paired spines have a greater extent over early stages development than in adulthood. Among amphibians, unpaired amphibians, in the form of a fold of skin devoid of a skeleton, are present as permanent or temporary formations in most larvae living in water, as well as in adult caudate amphibians and the larvae of tailless amphibians. Among mammals, P. are found in cetaceans and lilacs that have switched to an aquatic lifestyle for the second time. Gypsy cetaceans (vertical dorsal and horizontal caudal) and lilacs (horizontal caudal) do not have a skeleton; these are secondary formations that are not homologous (see Homology) to the unpaired P. of fish. Paired P. of cetaceans and lilacs, represented only by the anterior P. (the hind ones are reduced), have internal skeleton and are homologous to the forelimbs of all other vertebrates.

Lit. Guide to Zoology, vol. 2, M.-L., 1940; Shmalgauzen I.I., Fundamentals of comparative anatomy of vertebrate animals, 4th ed., M., 1947; Suvorov E.K., Fundamentals of Ichthyology, 2nd ed., M., 1947; Dogel V.A., Zoology of invertebrates, 5th ed., M., 1959; Aleev Yu. G., Functional principles of the external structure of fish, M., 1963.

V. N. Nikitin.

Big Soviet encyclopedia. - M.: Soviet Encyclopedia. 1969-1978 .

See what “Fins” are in other dictionaries:

- (pterigiae, pinnae), organs of movement or regulation of body position of aquatic animals. Among invertebrates, pelagics have P. forms of certain mollusks (modified leg or fold of skin), bristle-jawed. In skullless fish and larvae of fish, the unpaired P.... ... Biological encyclopedic dictionary

Organs of movement or regulation of body position of aquatic animals (some mollusks, chaetognaths, lancelets, cyclostomes, fish, some amphibians and mammals, cetaceans and sirenids). They can be paired or unpaired. * * * FINS… … encyclopedic Dictionary

Organs of movement or regulation of body position of aquatic animals (some mollusks, chaetognaths, lancelets, cyclostomes, fish, some amphibians and mammals, cetaceans and sirenids). There are paired and unpaired fins... Big Encyclopedic Dictionary

Fish fins can be paired or unpaired. The paired ones include the thoracic P (pinna pectoralis) and the abdominal V (pinna ventralis); to the unpaired ones - dorsal D (pinna dorsalis), anal A (pinna analis) and caudal C (pinna caudalis). The exoskeleton of the fins of bony fishes consists of rays that can be branchy And unbranched. Top part branched rays is divided into separate rays and has the appearance of a brush (branched). They are soft and located closer to the caudal end of the fin. Unbranched rays lie closer to the anterior edge of the fin and can be divided into two groups: articulated and non-articulated (spiny). Articulated the rays are divided along their length into separate segments; they are soft and can bend. Unarticulated– hard, with a sharp apex, tough, can be smooth or jagged (Fig. 10).

Figure 10 – Fin rays:

1 – unbranched, segmented; 2 – branched; 3 – prickly smooth; 4 – prickly jagged.

The number of branched and unbranched rays in the fins, especially in unpaired ones, is an important systematic feature. The rays are calculated and their number is recorded. Non-segmented (spiny) ones are designated by Roman numerals, branched ones - by Arabic numerals. Based on the calculation of the rays, a fin formula is compiled. So, pike perch has two dorsal fins. The first of them has 13-15 spiny rays (in different individuals), the second has 1-3 spines and 19-23 branched rays. The formula for the dorsal fin of pike perch is as follows: D XIII-XV, I-III 19-23. In the anal fin of pike perch, the number of spiny rays is I-III, branched 11-14. The formula for the anal fin of pike perch looks like this: A II-III 11-14.

Paired fins. All real fish have these fins. Their absence, for example, in moray eels (Muraenidae) is a secondary phenomenon, the result of late loss. Cyclostomes (Cyclostomata) do not have paired fins. This is a primary phenomenon.

The pectoral fins are located behind the gill slits of fish. In sharks and sturgeon, the pectoral fins are located in a horizontal plane and are inactive. These fish have a convex dorsal surface and a flattened ventral side of the body that gives them a resemblance to the profile of an airplane wing and creates lift when moving. Such an asymmetry of the body causes the appearance of a torque that tends to turn the fish’s head down. Pectoral fins and rostrum of sharks and sturgeon fish functionally they constitute a single system: directed at a small (8-10°) angle to the movement, they create additional lifting force and neutralize the effect of torque (Fig. 11). If a shark's pectoral fins are removed, it will raise its head upward to keep its body horizontal. In sturgeon fish, the removal of pectoral fins is not compensated for in any way due to poor flexibility of the body in the vertical direction, which is hampered by bugs, therefore, when the pectoral fins are amputated, the fish sinks to the bottom and cannot rise. Since the pectoral fins and rostrum in sharks and sturgeons are functionally connected, the strong development of the rostrum is usually accompanied by a decrease in the size of the pectoral fins and their removal from the anterior part of the body. This is clearly noticeable in the hammerhead shark (Sphyrna) and sawnose shark (Pristiophorus), whose rostrum is highly developed and the pectoral fins are small, while in the sea fox shark (Alopiias) and the blue shark (Prionace), the pectoral fins are well developed and the rostrum is small.

Figure 11 – Scheme of vertical forces arising during forward movement shark or sturgeon in the direction of the longitudinal axis of the body:

1 - center of gravity; 2 – center of dynamic pressure; 3 – force of residual mass; V 0 – lift force created by the body; V R– lifting force created by the pectoral fins; V r– lifting force created by the rostrum; Vv– lifting force created by the pelvic fins; V With– lift force created by the caudal fin; Curved arrows show the effect of torque.

The pectoral fins of bony fish, unlike the fins of sharks and sturgeons, are located vertically and can perform rowing movements back and forth. The main function of the pectoral fins of bony fishes is low-speed propulsion, allowing precise maneuvering when searching for food. The pectoral fins, together with the pelvic and caudal fins, allow the fish to maintain balance when motionless. The pectoral fins of stingrays, which evenly border their body, serve as the main propellers when swimming.

The pectoral fins of fish are very diverse in both shape and size (Fig. 12). In flying fish, the length of the rays can be up to 81% of the body length, which allows

Figure 12 – Shapes of pectoral fins of fish:

1 - flying fish; 2 – slider perch; 3 – keel belly; 4 – body; 5 – sea rooster; 6 - angler.

fish soar in the air. In freshwater fish, keelbellies from the Characin family, enlarged pectoral fins allow the fish to fly, reminiscent of the flight of birds. In gurnards (Trigla), the first three rays of the pectoral fins have turned into finger-like outgrowths, relying on which the fish can move along the bottom. Representatives of the order Anglerfish (Lophiiformes) have pectoral fins with fleshy bases that are also adapted to move along the ground and quickly bury themselves in it. Moving along hard substrates with the help of pectoral fins made these fins very mobile. When moving along the ground, anglerfish can rely on both pectoral and ventral fins. In catfish of the genus Clarias and blennies of the genus Blennius, the pectoral fins serve as additional supports during serpentine movements of the body while moving along the bottom. The pectoral fins of jumpers (Periophthalmidae) are arranged in a unique way. Their bases are equipped with special muscles that allow the fin to move forward and backward, and have a bend reminiscent of the elbow joint; The fin itself is located at an angle to the base. Living on coastal shallows, jumpers with the help of pectoral fins are able not only to move on land, but also to climb up plant stems, using the caudal fin with which they clasp the stem. With the help of pectoral fins, slider fish (Anabas) also move on land. Pushing off with their tail and clinging to plant stems with their pectoral fins and gill cover spines, these fish are able to travel from body of water to body of water, crawling hundreds of meters. In such benthic fish as rock perches (Serranidae), sticklebacks (Gasterosteidae), and wrasse (Labridae), the pectoral fins are usually wide, rounded, and fan-shaped. When they work, undulation waves move vertically downward, the fish appears to be suspended in the water column and can rise upward like a helicopter. Fishes of the order Pufferfish (Tetraodontiformes), pipefish (Syngnathidae) and pipits (Hyppocampus), which have small gill slits (the gill cover is hidden under the skin), can make circular movements with their pectoral fins, creating an outflow of water from the gills. When the pectoral fins are amputated, these fish suffocate.

The pelvic fins perform mainly the function of balance and therefore, as a rule, are located near the center of gravity of the fish's body. Their position changes with the change in the center of gravity (Fig. 13). In low-organized fish (herring-like, carp-like) the pelvic fins are located on the belly behind the pectoral fins, occupying abdominal position. The center of gravity of these fish is on the belly, which is due to the non-compact position of the internal organs occupying a large cavity. In highly organized fish, the pelvic fins are located in the front of the body. This position of the pelvic fins is called thoracic and is characteristic primarily of most perciform fish.

The pelvic fins can be located in front of the pectoral fins - on the throat. This arrangement is called jugular, and it is typical for large-headed fish with a compact arrangement of internal organs. The jugular position of the pelvic fins is characteristic of all fish of the order Codfish, as well as large-headed fish of the order Perciformes: stargazers (Uranoscopidae), nototheniids (Nototheniidae), blennies (Blenniidae), etc. Pelvic fins are absent in fish with eel-shaped and ribbon-shaped bodies. In erroneous (Ophidioidei) fish, which have a ribbon-eel-shaped body, the pelvic fins are located on the chin and serve as organs of touch.

Figure 13 – Position of the ventral fins:

1 – abdominal; 2 – thoracic; 3 – jugular.

The pelvic fins can be modified. With their help, some fish attach to the ground (Fig. 14), forming either a suction funnel (gobies) or a suction disk (lumpfish, slugs). The ventral fins of sticklebacks, modified into spines, have a protective function, and in triggerfishes, the pelvic fins have the appearance of a spiny spine and, together with the spiny ray of the dorsal fin, are a protective organ. In males cartilaginous fish the last rays of the pelvic fins are transformed into pterygopodia - copulatory organs. In sharks and sturgeons, the pelvic fins, like the pectoral fins, serve as load-bearing planes, but their role is less than that of the pectoral fins, since they serve to increase lifting force.

Figure 14 - Modification of the pelvic fins:

1 – suction funnel in gobies; 2 - suction disk of a slug.

Habitats and external structure of fish

The habitat of fish is various bodies of water on our planet: oceans, seas, rivers, lakes, ponds. It is very vast: the area occupied by the oceans exceeds 70% of the Earth’s surface, and the most deep depressions They go 11 thousand meters deep into the oceans.

The variety of living conditions in water influenced the appearance of fish and contributed to a wide variety of body shapes: the emergence of many adaptations to living conditions, both in structure and in biological characteristics.

General plan of the external structure of fish

On the head of the fish there are eyes, nostrils, a mouth with lips, and gill covers. The head smoothly transitions into the body. The body continues from the gill covers to the anal fin. The body of the fish ends with a tail.

The outside of the body is covered with skin. Protects mucus-coated skin of most fish scales .

The locomotion organs of fish are fins . Fins are outgrowths of skin resting on bones. fin rays . The caudal fin is of greatest importance. On the lower sides of the body there are paired fins: pectoral and ventral. They correspond to the fore and hind limbs of terrestrial vertebrates. The position of paired fins varies among different fish. The dorsal fin is located on top of the fish’s body, and the anal fin is located below, closer to the tail. The number of dorsal and anal fins may vary.

On the sides of the body of most fish there is a kind of organ that senses the flow of water. This lateral line . Thanks to the lateral line, even blinded fish do not bump into obstacles and are able to catch moving prey. The visible part of the lateral line is formed by scales with holes. Through them, water penetrates into a channel stretching along the body, to which the endings of nerve cells approach. The lateral line may be intermittent, continuous, or completely absent.

Functions of fins

Thanks to fins, fish are able to move and maintain balance in aquatic environment. Deprived of fins, it turns over with its belly up, since the center of gravity is located in the dorsal part.

Unpaired fins (dorsal and anal) provide stability to the body. The caudal fin in the vast majority of fish performs the function of propulsion.

Paired fins (thoracic and abdominal) serve as stabilizers, i.e. provide a balanced position of the body when it is immobile. With their help, the fish maintains its body in the desired position. When moving, they serve as load-bearing planes and steering wheels. The pectoral fins move the fish's body when swimming slowly. The pelvic fins perform mainly a balancing function.

Fish have a streamlined body shape. It reflects the characteristics of the environment and lifestyle. In fish adapted to fast, long-term swimming in the water column ( tuna(2), mackerel, herring, cod, salmon ), “torpedo-shaped” body shape. In predators that practice quick throws at short distances ( pike, taimen, barracuda, garfish (1) , saury), it is “arrow-shaped”. Some fish adapted to long-term residence on the bottom ( stingray (6) , flounder (3) ), have a flat body. In some species, the body has a bizarre shape. For example, sea Horse resembles a corresponding chess piece: its head is located at right angles to the axis of the body.

Sea Horses inhabit different oceans Globe. These fish surprise everyone who observes them: the body, like an insect, is enclosed in a shell, the prehensile tail of a monkey, the rotating eyes of a chameleon and, finally, a pouch like a kangaroo.

Although this cute fish can swim upright using the oscillatory movement of its dorsal fin, it is a poor swimmer and spends most of its time hanging, clinging to the seaweed with its tail and looking for small prey. The tubular snout of the skate acts like a pipette - when the cheeks are sharply inflated, the prey is quickly drawn into the mouth from a distance of up to 4 cm.

The smallest fish is considered Philippine bull Pandaku . Its length is about 7 mm. At one time fashionistas wore these fish in their ears. In crystal aquarium earrings!

Most big fish consider whale shark, which reaches a length of 15 m.

Additional fish organs

Some fish species (such as carp and catfish) have antennae around their mouths. This - additional organs touch and taste of food. In many deep-sea fish (for example, deep-sea anglerfish, hatchet fish, anchovy, photoblepharon ) luminous organs are developed.

There are protective spines on the scales of fish. They can be located in different parts bodies. For example, spines cover the body hedgehog fish .

Some fish, for example scorpionfish, sea dragon, wart They have organs of defense and attack - poisonous glands located at the base of the spines and fin rays.

Coverings of the body

On the outside, the skin of fish is covered with scales - thin translucent plates. The scales overlap each other with their ends, arranged in a tile-like manner. This provides

strong protection of the body and at the same time does not create obstacles to movement. Scales are formed by special skin cells. The size of the scales varies: from microscopic to blackheads up to several centimeters Indian barbel . There is a wide variety of scales: in shape, strength, composition, quantity and some other characteristics.

Lie in the skin pigment cells - chromatophores : when they expand, the pigment grains spread over a larger space and the color of the body becomes bright. If the chromatophores contract, the pigment grains accumulate in the center, leaving most of the cell uncolored, and the body color fades. If pigment grains of all colors are evenly distributed inside the chromatophores, the fish is brightly colored; if pigment grains are collected in the centers of cells, the fish becomes almost colorless and transparent; if only yellow pigment grains are distributed among their chromatophores, the fish changes color to light yellow.

Chromatophores determine the diversity of fish colors, which are especially bright in the tropics. Thus, fish skin performs the function of external protection. It protects the body from mechanical damage, facilitates sliding, determines the color of the fish, and communicates with external environment. The skin contains organs that sense temperature and chemical composition water.

Color meaning

Pelagic fish often have a dark "back" and a light "belly" like this fish abadejo cod family.

Indian glass catfish can serve as a textbook for studying anatomy.

Many fish that live in the upper and middle layers of water have a darker color on the upper part of the body and a lighter color on the lower part. The silvery belly of the fish, if you look at the fish from below, will not stand out against the light background of the sky. In the same way, the dark back, if you look at the fish from above, will merge with the dark background of the bottom.

By studying the coloration of fish, you can see how it helps camouflage and imitate other species of organisms, observe the demonstration of danger and inedibility, as well as the presentation of other signals by fish.

During certain periods of life, many fish acquire bright mating colors. Often the color and shape of the fish complement each other.

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The hydrosphere is characterized by an extreme diversity of conditions. These are fresh, flowing and standing waters, as well as salty seas and oceans inhabited by organisms at different depths. To exist in such diverse conditions, fish have developed both general principles of structure that meet the requirements of the environment (smooth, elongated body without protrusions, covered with mucus and scales; pointed head with pressed gill covers; fin system; lateral line), and adaptations characteristic of individual groups (flattened body, light organs, etc.). Each species of fish has numerous and varied adaptations corresponding to a specific way of life.

Pisces use many different methods to communicate. Of course, not as much as humans or other higher vertebrates. To communicate certain information to surrounding fish or other animals, fish can use chemical, electrolocation, sound and, as it turned out, visual methods, that is, they use “sign language” for communication. And although fishermen, unlike aquarists, divers or underwater hunters, are less likely to look a live fish in the eyes, some basic fish language can be learned.

Familiarization
The visible signals that fish can give to fish or other animals around them can be divided into several main groups. The first group is spawning postures or even gestures and facial expressions. After all, the movements of the fins can be called gestures, and the slightly open and even curved mouth can be called facial expressions.

The second group of visual signals demonstrates aggression, attack, and they indicate that this individual is “on the warpath.” There is also large group defensive gestures. This is not open aggression, but such gestures clearly show that we are peaceful fish, but “our armored train is on a siding.” Fish demonstrate these gestures more often than others.

The same group of gestures applies to protecting the territory, and to protecting one’s found (caught) food object, and to protecting the cubs.

Another important visual stimulus is the color of the fish. In a sufficient number of fish species, under stress, during spawning, during an aggressive attack or defense of their “good,” a color change occurs, which signals something out of the ordinary. Something similar happens to a person when, out of anger, shame or tension, he blushes and thereby betrays himself.

Unfortunately, the sign language of fish has not yet been fully studied and by no means for all species, but still knowledge general principles Gestural communication of fish will help to understand the fish. By the way, scientists suggest that fish of each species have a personal sign language, which is understood very well by closely related species and much worse by species that are far apart in their position in the taxonomy.

Gestures of aggression and defense
These gestures may, of course, vary among fish of different species, but they have much in common and are understandable to other fish. Greatest Animal Behavior Researcher, Laureate Nobel Prize Konrad Lorenz said: “Aggression is one of the most important factors in maintaining the structure of communities of most animal groups.”

Lorenz pointed out that the existence of groups with close individual connections between individuals is possible only in animals with sufficient developed ability to directed aggression, in which the combination of two or more individuals contributes to better survival.

In fish, the key aggressive gesture can be considered this: one of the fish turns to the other and begins to open its mouth wide (this is how dogs, wolves and other land animals grin). This gesture can be deciphered as a gesture of a frontal threat (attack).

So if a shark grins at you, leave quickly. While the mouth is just opening, this is some kind of beginning of a threat, territorial defense or any defensive gesture.

An important key point not only of this aggressive gesture, but also of other gestures of the same group: a fish with an open mouth seems larger, and therefore scarier and more impressive. At the same time, her attack looks more convincing and effective.

By the way, spreading the pectoral fins to the sides, protruding gill covers, and inflating the body with various tetraodons also leads to a general increase in the body volume of the frightening fish.

Male fish use certain poses of aggression and active defense to conquer females before spawning. There is no talk of direct use of gestures at this moment, but the female sees how big and serious the suitor is in front of her.

These “exaggeration” poses are very important for fish. After all, they grow throughout their lives, and for them size plays a primary role. Adult individuals, already showing aggressive behavior with all their might, are often large in size.

And the one who is larger is stronger, and older, and more experienced, and more important. That is, he has the right to food, territory, and the best female. Therefore, fish often try to visually exaggerate their size.

An exaggeration of size that frightens the enemy is also achieved by occupying a higher point in space. It is enough to force your opponent to look up, and he will feel inferior to you. Demonstration of the sides of the body and fluttering of the caudal fin and the entire body is often a manifestation of spawning behavior, that is, spawning gestures, or releasers.

However, in some fish (for example, ruffes and other perches), such a display of the sides and trembling of the tail is a typical aggressive gesture. This gesture of some fish is called a “side threat”. Unlike the “frontal threat”, it does not look so intimidating.

The spreading of the fins, often accompanied by trembling (or fluttering, or even shaking of the body), can be interpreted, depending on the situation, as aggression, as active defense, and as gestures of spawning behavior.

And in many territorial fish, such lateral displays, which are accompanied by vibrations of the body and spreading of the fins, have a dual function. For fish of the same species, but of the opposite sex, this is an attractive maneuver, showing what a beautiful, large and wonderful partner is swimming nearby.

And for relatives of the same sex, these gestures mean one thing: this is my female and my place, and you can leave! If one male (or female) spreads his fins, and his opponent, on the contrary, folds them, this means the latter’s complete surrender.

When the enemy in response inflates his fins and vibrates his body, this means that he is accepting the battle and there will be a show. A very important evolutionary point is the demonstration of aggression instead of a direct attack. Indeed, in its original form, aggression involves attacking an object, causing physical damage to it, or even killing it.

In the process of animal evolution, an aggressive attack was replaced by a demonstration of the threat of the possibility of an attack, especially during clashes between individuals of the same species. A demonstration, by causing fear in the enemy, allows you to win a skirmish without resorting to a fight, which is very dangerous for both sides.

Physical confrontation is replaced by psychological confrontation. Therefore, developed aggressive behavior, including many threats and frightening actions, is useful for the species, and for well-armed species it is simply life-saving.

This is why Lorenz argued that well-formed aggressive behavior is one of the remarkable achievements of natural selection and is essentially humane.

In fish, one of the main weapons of demonstration (instead of attack) is spines in the fins, spiny gill covers or plaques on the body. That is, the easiest way to scare the enemy is by showing him the means of defense and attack that he has at his disposal. this type animals.

Therefore, the fish, threatening, spread their fins and raise their spines; many stand upright in the water, exposing them to the enemy.

The fight process in fish consists of five to six successive phases:

warning with taking the appropriate posture;
excitement of opponents, usually accompanied by a change in color;
bringing fish closer together and demonstrating a threat pose;
mutual blows with the tail and mouth;
retreat and defeat of one of the opponents.

There are also phases of breaks to relieve tension and to rest during the battle or demonstration of strength.

Coloring and body pattern like spawning releasers
There are a lot of such visual and identification signals. During spawning, when the fish has a special hormonal background, many species change color and pattern - this is a signal that it is ready to reproduce.

For reliability, chemical and other signals also work actively, so that the fish is not mistaken and the species continues to exist. In addition to spawning, coloring and pattern help fish during schooling: often stripes on the body serve as a visual stimulus, helping thousands of fish stay close and correctly positioned relative to each other.

Coloring makes it possible to recognize your relative or, conversely, an enemy and dangerous individual. Many fish, especially those in which visual cues play important role(pike, perch, pike perch and others), remember well external features“our own” and “foreign” fish. Often two or three “lessons” are enough for the fish to remember well the color and pattern of the hostile fish.

Sometimes not only the color of the entire body, but also the color of individual fins (for example, abdominal or pectoral), or individual brightly colored areas on the body (abdomen, back, head) signal to potential partners that “ready to spawn!”

A spot on the abdomen of many females indicates that there is a lot of caviar in the abdomen, it is enlarged and bright. However, in most cases, bright coloring is destructive outside of spawning: peaceful fish it unmasks itself in front of predators, and, on the contrary, reveals the predator ahead of time.

So most of the fish in our reservoirs during the normal non-spawning period have a gray, inconspicuous appearance, and developed gesticulation is all the more important for them.
In addition to spawning behavior or identification of “friend” or “alien,” coloration can work as a factor determining status.

The brighter the color and clearer the pattern, the higher the social status of this individual. This is not always the case, but it is often the case. Fish can use their coloration to demonstrate threat (strong, intense coloration) or submission (less bright or dull coloration), usually this is supported by appropriate gestures that reinforce the information. Bright coloring is actively used by fish that protect their offspring, raise young and drive away other fish that are dangerous to the young. It also helps juveniles identify their parents and notice them among other fish.

In parental behavior, fish have a highly developed not only body color language, but also body language. The young quickly remember that the flapping of the pelvic fins and the pressed pectoral fins mean the call “swim to mom”; a bend of the body and a slightly open mouth - “swim after me”; fins spread out are a command to hide in cover.

For normal relationships between parents and young, it is necessary to suppress certain reactions. Very interesting examples of this have been observed in fish. Some chromis (family Cichlids) carry young fish in their mouths; At this time, adult fish do not feed at all.

A funny case is described with a male of one species of chromis, whose representatives every evening transfer the young to the “bedroom” - a hole dug in the sand. This “father” was collecting fry in his mouth, grabbing the ones that had strayed to the side one by one, and suddenly he saw a worm: after hesitating a little, he finally spat out the fry, grabbed and swallowed the worm, and then again began to collect the “cubs” to transfer them to the hole .

A straightened, standing dorsal fin indicates the beginning aggressive behavior(for example, when protecting one’s territory), and about an invitation to spawn.

Rituals and demonstrations
To understand the sign language of fish, you need to know their rituals and the meaning of different poses and gestures, which say a lot about the fish’s intentions. Rituals and demonstrative acts of behavior exhibited by animals in conflict situations can be divided into two groups: rituals of threat and rituals of pacification, inhibiting aggression from stronger relatives. Lorenz identified several main features of such rituals.

Demonstrative exposure of the most vulnerable part of the body. It is very interesting that dominant animals often demonstrate this behavior. So, when two wolves or dogs meet, the stronger animal turns its head away and exposes to its opponent the area of the carotid artery, curved towards the bite.

The meaning of such a demonstration is that the dominant signals in this way: “I’m not afraid of you!” This is more likely to apply to more highly developed animals, but some fish also exhibit similar behavior. For example, cichlids show folded fins and caudal peduncle to a strong enemy.

Fish have organs that can be called organs of ritual behavior. These are fins and gill covers. Ritual ones are modified fins, which in the process of evolution turn into thorns or spines or, conversely, into veil formations. All these “decorations” are clearly demonstrated in front of other individuals of their species, in front of a female or a rival. Coloring can also be ritualistic.

For example, tropical fish have a false “eye” - a bright spot in the upper corner of the dorsal fin that imitates the eye of a fish. The fish exposes this corner of the fin to the enemy, the enemy grabs onto it, thinking that it is an eye and that he will now kill the victim.

And he just rips out several rays of the dorsal fin with this bright spot, and the victim swims away safely almost unharmed. Obviously, in the course of evolution, both the decorations themselves and the ways of displaying them developed in parallel.

The demonstration of signal structures carries vital information that indicates to other individuals the gender of the demonstrating animal, its age, strength, ownership of a given area of the area, etc.

Ritual demonstrations of territorial behavior among fish are very important and interesting. The forms of aggressive territorial behavior themselves are far from being limited to direct attacks, fights, chases, etc. One can even say that such “harsh” forms of aggression, associated with inflicting wounds and other damage to the enemy, are not such a common phenomenon in the general system of individualization of territory.

Direct aggression is almost always accompanied by special “ritual” forms of behavior, and sometimes the protection of an area is completely limited to them. And the clashes themselves on territorial grounds are relatively rarely accompanied by causing serious damage to the enemy. Thus, frequent fights of goby fish at the boundaries of areas are usually very short-lived and end with the flight of the “intruder”, after which the “owner” begins to swim vigorously in the reclaimed area.

Fish actively mark their territory. Each species does this in its own way, depending on which sensory systems prevail in a given species. Thus, the territory is visually marked by species that live in small, easily visible areas. For example, the same coral fish. A clear, bright, unusual and different from other fish body pattern (and coloring) - all this in itself indicates that the owner of the population of this species is located in this area.

Hierarchy and poses of fish with gestures
The first meeting of animals rarely occurs without some tension, without mutual manifestation of aggressiveness. A fight breaks out, or the individuals demonstrate their unfriendliness with decisive gestures and threatening sounds. However, after the relationship is cleared up, fights rarely occur. When meeting again, animals unquestioningly give way to a stronger opponent the road, food or other object of competition.

The order of subordination of animals in a group is called hierarchy. Such orderliness of relationships leads to a reduction in energy and mental costs that arise from constant competition and clarification of relationships. Animals at the lower levels of the hierarchy, subject to aggression from other members of the group, feel oppressed, which also causes important physiological changes in their body, in particular the occurrence of an increased stress response. It is these individuals who most often become victims natural selection.

Each individual is either superior in strength to its partner or inferior to it. Such a hierarchical system is formed when fish collide in the struggle for a place in a reservoir, for food and for a female.

The fish just opened its mouth and raised its fin, and its size visually increased by almost 25%. This is one of the most accessible and common ways to raise your authority in the animal world.

In the early stages of establishing a hierarchy, a lot of fights occur between fish (which are inherently hierarchical in principle). After the final establishment of the hierarchy, aggressive collisions between fish individuals practically cease, and the order of subordination of individuals is maintained in the population.

Usually, when a high-ranking fish approaches, subordinate individuals yield to it without resistance. In fish, size is most often the main criterion for dominance in the hierarchical ladder.
The number of collisions in a group of animals increases sharply when there is a lack of food, space or other living conditions. Lack of food, causing more frequent collisions of fish in a school, causes them to spread out somewhat to the sides and take over additional feeding area.

Fatal fights of very aggressive fish species in fish farms and aquariums are observed much more often than in natural conditions. This can easily be explained by stress and the inability to separate opponents. A kind of eternal ring. Therefore, aquarists know how important it is to provide plenty of hiding places in a pond if the fish are territorial. It’s even safer to keep them separate.

The lower links of the fish in the hierarchical ladder should demonstrate postures of subordination, humility and appeasement. What does a losing fish do? First of all, she raises the “white flag”, that is, folds her fins, removes thorns, thorns and teeth (sharks). These attributes of aggressiveness are removed until better times, that is, before meeting an even weaker opponent.

The size of individuals decreases before our eyes. As far as possible, of course. That is, the losing outsider fish demonstrates to the enemy: “I’m small and unarmed, I’m not afraid of you!” And the strong, victorious opponent also understands that he no longer needs to demonstrate his strength, and closes his mouth, assumes a horizontal position, folds his fins, removes thorns and thorns (if there are any, of course).

Sometimes a defeated fish turns over with its belly up and this also demonstrates its defenselessness. I deliberately do not present data on specific species here, since there are very few of them, and many have not yet been statistically confirmed.

I hope that interesting information will help anglers better understand fish, and not once again scare or cause harm to either a specific fish or a school or population as a whole.

Source: Ekaterina Nikolaeva, Fish with us 3/2013 159

Gustera

Silver bream fish. The silver bream differs from the above-described species of bream solely in the number and location of the pharyngeal teeth, of which there are not five, but seven on each side, and moreover, in two rows. In body shape it is very similar to a young bream or, more accurately, a bream, but has a smaller number of rays in the dorsal (3 simple and 8 branched) and anal (3 simple and 20-24 branched) fins; in addition, its scales are noticeably larger, and its paired fins are reddish in color.

The body of the silver bream is strongly flattened, and its height is at least a third of its entire length; her nose is blunt, her eyes are large and silvery; the back is bluish-gray, the sides of the body are bluish-silver; unpaired fins are gray, and paired fins are red or reddish at the base, dark gray towards the apex. However, this fish, depending on age, time of year and local conditions, presents significant modifications.

Gustera never reaches a significant size. For the most part it is no more than one pound and less than a foot in length; One and a half and two pound ones are less common, and only in a few places, for example in the Gulf of Finland. Lake Ladoga, it weighs up to three pounds. This fish has a much wider distribution than syrty, bluefish and glazach.

Gustera is found in almost all European countries: France, England, Sweden, Norway, throughout Germany, Switzerland, and it seems to be absent only in Southern Europe. In all the above-mentioned areas, it belongs to very ordinary fish. In Russia, the silver bream is found in all rivers, sometimes even small rivers, also in lakes, especially in the northwestern provinces, and flowing ponds; in Finland it reaches 62° N. sh.; is also found in the northern parts of Lake Onega, and in northern Russia goes even further - to Arkhangelsk.

It seems that it no longer exists in Pechora, and in Siberia it was found only recently (Varpakhovsky) in the river. Iset, a tributary of the Tobol. There is no silver bream in the Turkestan region, but in Transcaucasia it has so far been found at the mouth of the Kura River and in Lake. Paleostome, off the coast of the Black Sea. Silver bream is a sluggish, lazy fish and, like bream, loves quiet, deep, fairly warm water, with a silty or clayey bottom, which is why it is very often found with this latter.

It lives in one place for a long time and most willingly stays near the very shores (hence its French name - la Bordeliere and Russian berezhnik), especially in the wind, since the shafts, eroding the banks, and in shallow places the very bottom, reveal various worms and larvae. In no large quantities it apparently lives at the mouths of rivers and at the seaside itself, as, for example, at the mouths of the Volga and in the Gulf of Finland between St. Petersburg and Kronstadt.

In spring and autumn, the silver bream is found in extremely dense flocks, which, of course, is where its common name comes from. However, it rarely makes very long journeys and almost never reaches, for example, the middle reaches of the Volga, where its own local bream lives. In general, the main mass of these fish accumulate in the lower reaches of rivers, in the sea, and, like very many others, they make regular periodic movements: in the spring they go up for spawning, in the fall for wintering.

Entering wintering grounds in the fall, they lie down in holes under the riffles in such large masses that in the lower reaches of the Volga it happens that up to 30 thousand of them can be pulled out in one ton. The food of the silver bream is almost the same as that of other types of bream: it feeds exclusively on mud and small mollusks, crustaceans and worms contained in it, most often bloodworms, but it also destroys the eggs of other fish, especially (according to Bloch’s observations) rudd caviar.

Spawning of the silver bream begins very late, b. hours after the end of bream spawning - at the end of May or at the beginning of June, in the south a little earlier. At this time, its scales change color, and the paired fins acquire a brighter red color; in males, in addition, small granular tubercles develop on the gill covers and along the edges of the scales, which then disappear again. Usually, small silver bream spawns earlier, large ones later.

In the Gulf of Finland, other fishermen distinguish two breeds of silver bream: one breed, according to them, is smaller, lighter in color, spawns earlier and is called Trinity (based on the time of spawning), and the other breed is much larger (up to 3 pounds), darker in color, spawns later and is called Ivanovskaya. According to Bloch’s observations, in Germany the largest silver bream spawns first, followed by the smallest one a week or nine days later.

The silver bream chooses grassy and shallow bays as a spawning site and spawns eggs extremely noisily, like bream, but incomparably quieter than it: at this time it sometimes even happens to catch them with your hands; then they catch her in the muzzle, winged and nonsense by the pounds. It usually spawns from sunset to ten o'clock in the morning, and each age finishes the game at 3-4 at night, but if cold weather interferes, then in one day.

In a medium-sized female, Bloch counted more than 100 thousand eggs. According to Sieboldt, the silver bream becomes capable of reproduction very early, not yet reaching 5 inches in length, so we must assume that it spawns in its second year. The main catch of silver bream is carried out in the spring - with seines, but in the lower reaches of rivers, especially on the Volga, even greater catches of this fish occur in the fall. The most full information about the crucian fish - here.

Silver bream generally belongs to low-value fish and is rarely prepared for future use, unless it is caught in very large quantities. Salted and dried silver bream on the lower Volga is sold under the name tarani; in the rest of the Volga region she b. h. is sold fresh and has only local sales. However, it is very suitable for fish soup and is held in rather greater esteem in the Volga provinces, where there is a saying about it: “Large silver bream is tastier than small bream.”

Where there is a lot of silver bream, it takes the bait very well, especially after spawning. In some places they usually fish for silver bream with a worm, from the bottom, like bream, and its bite is similar to the latter’s bite; The silver bream, even more often than the bream, drags the float to the side without submerging it, and often hooks itself. This is perhaps the most daring and annoying fish, which is pure punishment for anglers fishing with bait.

It has been noticed that she takes best at night. According to Pospelov, the silver bream on the river. Teze (in Vladimir province) is caught as if with pieces of salted herring. In Germany in the fall it also goes well for bread with honey, and on the Volga it is very often caught in winter from ice holes (using a worm). The winter bite of the silver bream has the usual character - it first twitches, then slightly drowns. For catching catfish, pike and large perch, silver bream is one of the the best attachments, since it is much more tenacious than other types of bream.

In many places in Russia, for example. in the Dnieper, Dniester, on the middle and lower Volga, occasionally - usually alone and in schools of other fish, b. including silver bream and roach (roach) - there is one fish that occupies, as it were, the middle between bream, silver bream and roach (Abramidopsis), on the river. In Mologa this fish is called ryapusa, in Nizhny Novgorod, Kazan and on the Dnieper - all fish, all fish, on the grounds that they resemble different carp fish: blue bream, silver bream, roach, rudd.

According to fishermen, as well as some scientists, this is a bastard from bream and roach or silver bream and roach. In Kazan, one fisherman even claimed to prof. Kessler that all fish hatch from roach eggs fertilized by male silver bream. In terms of body shape and pharyngeal teeth, this cross is still closer to the genus Abramis.

The height of its body is about 2/7 of the entire length, the mouth occupies the top of the snout and the lower jaw is slightly turned upward; the scales are larger than those of other breams, and the anal fin contains only 15-18 unbranched rays; the lower lobe of the caudal fin is barely longer than the upper one, before Abramidopsis is already approaching the roach. It would be more accurate to assume that this for the most part a cross between bream and roach.

A similar cross is Bliccopsis abramo-rutilus Holandre, which probably descended from silver bream and roach and was occasionally found here and there alone, both in central Europe and in Russia. According to Kessler, Bliccopsis is also found in lake. Paleostom (at the mouth of Rion in the Caucasus). The body of the silver bream is tall, strongly compressed laterally, covered with thick, tightly fitting scales. Her head is relatively small. The mouth is small, oblique, semi-inferior, retractable.

The eyes are big. The dorsal fin is high, the anal fin is long. The back is bluish-gray, the sides and belly are silver. The dorsal, caudal and anal fins are gray, the pectoral and ventral fins are yellowish, sometimes reddish, which is how it differs in appearance from bream. In addition, the silver bream, unlike the bream, has larger scales, especially at the dorsal fin, as well as on the back; behind the back of the head it has a groove not covered with scales.

The silver bream lives in rivers, lakes and ponds. In rivers, it sticks to places with a slow flow and considerable depth, as well as in bays, backwaters, oxbow lakes, where the bottom is sandy and clay with a small admixture of silt. It is most numerous in lakes and in lowland areas of rivers. Large individuals live in bottom layers of water, deep pools, holes and in open areas of lakes and reservoirs.

The smaller silver bream prefers to stay in coastal areas among sparse thickets. At the same time, small individuals usually stay in large flocks. Gustera is characterized by a sedentary lifestyle. In summer its flocks are small. With the onset of autumn cold weather, they increase in size and move to the pits. With the onset of spring floods, its flocks go to feeding areas.

As spawning time approaches, after the water warms up, flocks of silver bream increase and move to the spawning grounds. At the same time, the lake spawning silver bream goes to the shores in large numbers, and the river bream, leaving the channel, enters small bays and creeks. The silver bream spawns from the end of April to May at a water temperature of 12-20°. During prolonged cold spells, spawning may last until June.

The white bream spawns in portions, but there are females that spawn at once. Its spawning occurs amicably, mainly in the evening and morning with a short night break. Before spawning, they become bright silver, the pectoral and ventral fins acquire an orange tint. Lumps of pearly rash appear on the head and upper body of spawning males. Soon after spawning, all mating changes disappear.

In the Dnieper, on the site of the now existing Kiev reservoir, three-year-old females of the silver bream had an average of 9.5 thousand eggs, six-year-olds - 22 thousand, and three years after the formation of the reservoir, more than 16 thousand eggs were found in three-year-old females, in six-year-olds - more than 80 thousand pieces, i.e., in the conditions of the reservoir, its fertility increased by 2-3 times.

The silver bream becomes sexually mature at two or three years of age, and in the spawning herd, males mature predominantly earlier than females. In older age groups The spawning stock of males is significantly smaller than females. The silver bream grows slowly. For example, in the lower reaches of the Southern Bug, yearlings had an average body length of 3.3 cm, three-yearlings - 10.2 cm, six-yearlings - 16.9 cm.

Until puberty, both sexes grow equally, but after puberty, the growth of males slows down somewhat. Juvenile silver bream in Dnieper reservoirs feed on crustaceans and chironomid larvae. To a lesser extent, it consumes algae, caddisflies, spiders and water bugs. Adult fish feed on higher aquatic plants, worms, mollusks, crustaceans, larvae and pupae of mosquitoes and other insects.

The main feeding grounds for small silver bream (10-15 cm long) are located mainly in the coastal zone. Large fish, feeding mainly on mollusks, feed in places more distant from the shore. Fish with a length of 25-32 cm, which have significant fatty deposits in their intestines, feed weaker. As the body size of the silver bream increases, the number of crustaceans and insect larvae in its food decreases and the number of mollusks increases.

It switches to feeding on mollusks when its body length is 13-15 cm or more. Depending on the composition and development food base the ratio of food organisms in the food composition of fish of the same size is not the same. For example, fish 10-12 cm long in the coastal zone feed mainly on insect larvae, and in deeper places on crustaceans, which corresponds to the distribution of these organisms in reservoirs.

White bream is widespread in Europe. It is absent in the rivers of the Arctic Ocean and in Central Asia. In the CIS it lives in the basins of the Baltic, Black, Azov and Caspian seas. In Ukraine, it lives in the basins of all rivers, excluding the rivers of Crimea and mountainous sections of other rivers.

List of fish: whitefish species muksun, omul and vendace

There are many salmon fish, one of the families is whitefish, a numerous, little-studied fish genus with variable characteristics. Representatives of this family have a laterally compressed body and a small mouth for their size, which causes a lot of inconvenience for fans of fishing with a rod. The lip of a whitefish often cannot withstand the load when it is pulled out of the water, and when the lip breaks off, the fish leaves.

Due to the similarity of the silhouette of the whitefish's head with the head of a herring, whitefish are also called herring, and only the adipose fin clearly indicates their salmon origin. The extremely high degree of variability of characters still does not allow us to establish the exact number of their species: in each lake it is possible to establish its own special kind, for example, 43 forms were identified only in the lakes of the Kola Peninsula. Currently, work is underway to combine similar forms into one species, which should lead to the systematization of fish species of the whitefish family.

General description of the family

On the territory of Russia there are over a hundred varieties of fish of this family, which have excellent taste and other beneficial properties. Its habitat is almost all bodies of water from the Kola Peninsula in the west to the Kamchatka and Chukotka Peninsulas in the east. Although this fish belongs to the salmon family, its meat is white, sometimes pinkish in color. Often, even experienced fishermen do not even suspect that the Baikal omul is the same whitefish. Here is a small list of the names of fish of the whitefish family:

largemouth and European vendace (ripus), Atlantic and Baltic whitefish;
whitefish Volkhovsky, Bauntovsky and Siberian (Pyzhyan), Baikal omul;
Muksun, Tugun, Valaamka and Chir (Shokur).

This diverse fish does not have a single appearance, but all members of the family have uniform silvery scales and darkened fins. Adipose fin, a distinctive feature of all salmon fish is also common feature fish of the whitefish genus. Distinctive feature females - scales, unlike the scales of males, they are larger and have a yellowish tint.

Like salmon, whitefish can be found in both fresh and salt water. Depending on this, two groups of whitefish are distinguished:

freshwater – lake and river;
anadromous or sea whitefish.

Gallery: whitefish species (25 photos)

Habits and preferences

A quality common to the entire family is life in a flock, which is formed according to the age of the individuals. Whitefish preferences are uncomplicated cold water, enriched with oxygen, which usually occurs in rapids of rivers and in the depths of lakes. At the same time, a school of whitefish can drive representatives of other fish species out of the pit. As a rule, than bigger fish, the further it goes from the coast.

The ability to spawn in fish of the family appears at the age of about three years, and in some breeds – a year or two later. Spawning of sea and freshwater whitefish takes place in the same conditions - all of them, including lake ones, rise to the upper reaches of rivers and their tributaries. Whitefish lay eggs in the fall, when the water cools to below five degrees. The spawning areas are deep holes and quiet rivers and reaches. Here the eggs are aged until spring, when the fry emerge from the eggs as the water warms.

The diet of the whitefish family, like all predators, is of animal origin: vertebrate and invertebrate insects (worms, larvae and caterpillars, caddis flies and bark beetles), small crustaceans and mollusks, caviar. Depending on the age and, accordingly, the size of the predator itself, it also attacks fish that are smaller than it. But among whitefish there are also lovers of vegetarian food collected from the bottom, as well as omnivores - semi-predators.

Their lifespan is about two decades, but fish of half their age are often caught. The largest whitefish is usually a little more than half a meter long, and small adult breeds are from one to one and a half decimeters.

As a rule, whitefish are divided into separate groups based on mouth position. The mouth can be directed upward - the upper mouth, forward - the terminal mouth, and downward - the lower mouth.

Topmouth are small fish that feed on what they find near the surface of the water. These are insects and invertebrates - worms and caterpillars. The fish with the upper mouth are represented mainly by the European vendace (ripus) and the larger Siberian one. The latter can be up to half a meter in length, lives in places where rivers flow into salt waters sea, almost never found in lakes. The rhipus is half the size and is an inhabitant of lakes. Both species of vendace are commercially available.

Whitefish with a mouth in front (final) are also considered commercial fish. Omul is a large fish, over half a meter long, which, like vendace, lives in the bays of the seas and the estuaries of rivers flowing into the sea, where it rises to spawn. The diet of omul includes crustaceans and small fish. Baikal omul is a lake variety of whitefish. Another lake-river variety is peled fish (curd), in sea water it does not enter, but is as large as vendace and omul, its length is about half a meter. It was also brought into reservoirs Southern Urals, here its size is not so impressive. There is also a small relative of whitefish with a terminal mouth - tugun, which lives in the rivers of Siberia. Its length does not exceed twenty centimeters.

Whitefish with a lower mouth also live in Russian water bodies; there are seven species of them. But work is currently underway to separate them, and there is no point in providing any information on them.

Freshwater whitefish

The river whitefish breed - by name, is an inhabitant of rivers, where it comes from the sea or a large lake when moving to spawn. His usual weight is about a kilogram, rarely exceeds two kilograms. River whitefish only winter in lakes; at all other times of the year they lead a river life. Essentially, it is acclimatized to river life marine or migratory whitefish. The caviar of this species of whitefish is numerous - up to 50 thousand eggs and slightly lighter than trout caviar.

Pechora whitefish, the most famous are omul, it was already mentioned above, peled, whitefish. The peled reaches a length of more than half a meter and a weight of about three kilograms. Chir is much larger, it can weigh up to ten kg, and lives in the lakes of the Pechora River basin and its channels.

The Baikal omul reaches a weight of up to seven kilograms; its food is small epishura crustaceans, and if there are insufficient quantities of them, it switches to eating small fish. Starting in September, the omul rises into the rivers, preparing for spawning. Based on the location of the spawning grounds, subspecies of the Baikal omul are distinguished:

Angara - early ripening, maturity at five years, but with slow growth;
Selenga - maturity at seven years, grows quickly;
Chivyrkuisky - also grows quickly, spawning in October.

The omul finishes spawning when slush already appears on the river and floats back to Lake Baikal for the winter. At one time, the fish was intensively caught by commercial fishermen, and its numbers decreased significantly, but now measures are being taken to artificially reproduce the omul.

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